Journal of Integrative Plant Biology

Article	Citations
Contribution of phenylpropanoid metabolism to plant development and plant–environment interactions	691
Protein kinases in plant responses to drought, salt, and cold stress	332
The plant cell wall: Biosynthesis, construction, and functions	245
Mitogen‐activated protein kinase cascades in plant signaling	223
Potassium and phosphorus transport and signaling in plants	206
The root microbiome: Community assembly and its contributions to plant fitness	160
Hydrogen sulfide, a signaling molecule in plant stress responses	130
Root developmental responses to phosphorus nutrition	127
Plant Metabolic Network 15: A resource of genome‐wide metabolism databases for 126 plants and algae	125
Molecular mechanisms for the photoperiodic regulation of flowering in soybean	123
Mechanisms underlying legume–rhizobium symbioses	117
Auxin signaling: Research advances over the past 30 years	112
Ethylene signaling in rice and Arabidopsis: New regulators and mechanisms	111
Ca2+ signaling in plant responses to abiotic stresses	101
The calcium‐dependent protein kinase ZmCDPK7 functions in heat‐stress tolerance in maize	85
The direct targets of CBFs: In cold stress response and beyond	81
Genome editing for plant research and crop improvement	75
An update on the function and regulation of methylerythritol phosphate and mevalonate pathways and their evolutionary dynamics	75
Phyllosphere microbiota: Community dynamics and its interaction with plant hosts	74
Phylotranscriptomic insights into Asteraceae diversity, polyploidy, and morphological innovation	68
Molecular mechanisms underlying the toxicity and detoxification of trace metals and metalloids in plants	66
Increasing yield potential through manipulating of an ARE1 ortholog related to nitrogen use efficiency in wheat by CRISPR/Cas9	65
Plant abiotic stress: New insights into the factors that activate and modulate plant responses	64
A rare Waxy allele coordinately improves rice eating and cooking quality and grain transparency	63
Transcriptional regulation of fleshy fruit texture	61

Single‐cell RNA sequencing reveals a high‐resolution cell atlas of xylem in Populus	59
Effective methods for isolation and purification of extracellular vesicles from plants	58
Plant plasma membrane‐resident receptors: Surveillance for infections and coordination for growth and development	58
Arabidopsis U‐box E3 ubiquitin ligase PUB11 negatively regulates drought tolerance by degrading the receptor‐like protein kinases LRR1 and KIN7	58
Chromatin remodeling factors regulate environmental stress responses in plants	54
Ethylene and salicylic acid synergistically accelerate leaf senescence in Arabidopsis	54
Phylogenomics and the flowering plant tree of life	50
Advances in structure and function of auxin response factor in plants	49
Salicylic acid and ethylene coordinately promote leaf senescence	49
From genes to networks: The genetic control of leaf development	49
The origin of Wxla provides new insights into the improvement of grain quality in rice	49
ERF1 delays flowering through direct inhibition of FLOWERING LOCUS T expression in Arabidopsis	48
Phylogenomic conflict analyses in the apple genus Malus s.l. reveal widespread hybridization and allopolyploidy driving diversification, with insights into the complex biogeographic history in	48
Histone deacetylase HDA710 controls salt tolerance by regulating ABA signaling in rice	48
Rice seed storage proteins: Biosynthetic pathways and the effects of environmental factors	47
Functional assembly of root‐associated microbial consortia improves nutrient efficiency and yield in soybean	47
Efficient and genotype independent maize transformation using pollen transfected by DNA‐coated magnetic nanoparticles	47
Versatile effectors of phytopathogenic fungi target host immunity	46
Creation of aromatic maize by CRISPR/Cas	45
SIZ1 negatively regulates aluminum resistance by mediating the STOP1–ALMT1 pathway in Arabidopsis	44
Abscisic acid signaling negatively regulates nitrate uptake via phosphorylation of NRT1.1 by SnRK2s in Arabidopsis	43
FT5a interferes with the Dt1‐AP1 feedback loop to control flowering time and shoot determinacy in soybean	43
Chromatin‐remodeling complexes: Conserved and plant‐specific subunits in Arabidopsis	43
Maize endosperm development	43
The Class III peroxidase gene OsPrx30, transcriptionally modulated by the AT‐hook protein OsATH1, mediates rice bacterial blight‐induced ROS accumulation	42
Establishment of a dmp based maternal haploid induction system for polyploid Brassica napus and Nicotiana tabacum	40
Crop phenotyping in a context of global change: What to measure and how to do it	39
Drought stress and plant ecotype drive microbiome recruitment in switchgrass rhizosheath	39
Integration of light and temperature signaling pathways in plants	38
The NPR1‐WRKY46‐WRKY6 signaling cascade mediates probenazole/salicylic acid‐elicited leaf senescence in Arabidopsis thaliana	38
Engineering broad‐spectrum disease‐resistant rice by editing multiple susceptibility genes	37
Highly efficient, genotype‐independent transformation and gene editing in watermelon (Citrullus lanatus) using a chimeric ClGRF4‐GIF1 gene	37
The lncRNA39896–miR166b–HDZs module affects tomato resistance to Phytophthora infestans	37
Fig fruit ripening is regulated by the interaction between ethylene and abscisic acid	36
Large chromosomal segment deletions by CRISPR/LbCpf1‐mediated multiplex gene editing in soybean	36
A recent retrotransposon insertion of J caused E6 locus facilitating soybean adaptation into low latitude	36
Loss of Gn1a/OsCKX2 confers heavy‐panicle rice with excellent lodging resistance	36
Circadian clock in plants: Linking timing to fitness	36
Recent progression and future perspectives in cotton genomic breeding	35
The battle of crops against drought: Genetic dissection and improvement	35
Efficient generation of homozygous substitutions in rice in one generation utilizing an rABE8e base editor	35
The CBP/p300 histone acetyltransferases function as plant‐specific MEDIATOR subunits in Arabidopsis	35
WRKY33‐mediated indolic glucosinolate metabolic pathway confers resistance against Alternaria brassicicola in Arabidopsis and Brassica crops	35
Comprehensive phylogenetic analyses of Orchidaceae using nuclear genes and evolutionary insights into epiphytism	35
Development of an efficient plant dual cytosine and adenine editor	34
Regulation of cytoskeleton‐associated protein activities: Linking cellular signals to plant cytoskeletal function	34
Salicylic acid biosynthesis is not from phenylalanine in Arabidopsis	34
Variations in OsSPL10 confer drought tolerance by directly regulating OsNAC2 expression and ROS production in rice	33
Efficient genotype‐independent cotton genetic transformation and genome editing	33
Plant base editing and prime editing: The current status and future perspectives	33

The MIEL1‐ABI5/MYB30 regulatory module fine tunes abscisic acid signaling during seed germination	33
Salicylic acid promotes quiescent center cell division through ROS accumulation and down‐regulation of PLT1, PLT2, and WOX5	33
Rice FLOURY ENDOSPERM 18 encodes a pentatricopeptide repeat protein required for 5′ processing of mitochondrial nad5 messenger RNA and endosperm development	33
Plant target of rapamycin signaling network: Complexes, conservations, and specificities	32
Structure of plant photosystem I−light harvesting complex I supercomplex at 2.4 Å resolution	32
Allele‐aware chromosome‐scale assembly of the allopolyploid genome of hexaploid Ma bamboo (Dendrocalamus latiflorus Munro)	32
Shaping polyploid wheat for success: Origins, domestication, and the genetic improvement of agronomic traits	32
The jasmonate‐induced bHLH gene SlJIG functions in terpene biosynthesis and resistance to insects and fungus	32
MdbHLH3 modulates apple soluble sugar content by activating phosphofructokinase gene expression	32
Ethylene‐induced stomatal closure is mediated via MKK1/3–MPK3/6 cascade to EIN2 and EIN3	32
BioClay™ prolongs RNA interference‐mediated crop protection against Botrytis cinerea	31
The miR166 mediated regulatory module controls plant height by regulating gibberellic acid biosynthesis and catabolism in soybean	31
Alternative splicing of MaMYB16L regulates starch degradation in banana fruit during ripening	31
Water stress resilient cereal crops: Lessons from wild relatives	31
RNA silencing: From discovery and elucidation to application and perspectives	31
Nitrate transporter NRT1.1 and anion channel SLAH3 form a functional unit to regulate nitrate‐dependent alleviation of ammonium toxicity	31
MiR319‐targeted OsTCP21 and OsGAmyb regulate tillering and grain yield in rice	31
Multiplex and optimization of dCas9‐TV‐mediated gene activation in plants	30
A unique photosystem I reaction center from a chlorophyll d‐containing cyanobacterium Acaryochloris marina	30
CDE4 encodes a pentatricopeptide repeat protein involved in chloroplast RNA splicing and affects chloroplast development under low‐temperature conditions in rice	30
OsMKKK70 regulates grain size and leaf angle in rice through the OsMKK4‐OsMAPK6‐OsWRKY53 signaling pathway	30
Nicotiana benthamiana LRR‐RLP NbEIX2 mediates the perception of an EIX‐like protein from Verticillium dahliae	29
Ustilaginoidea virens modulates lysine 2‐hydroxyisobutyrylation in rice flowers during infection	29
Reactive oxygen species: Multidimensional regulators of plant adaptation to abiotic stress and development	29
CRISPR/Cas9 gene editing and natural variation analysis demonstrate the potential for HvARE1 in improvement of nitrogen use efficiency in barley	29
Here comes the sun: How optimization of photosynthetic light reactions can boost crop yields	29
GmPIN1‐mediated auxin asymmetry regulates leaf petiole angle and plant architecture in soybean	28
Integrated view of plant metabolic defense with particular focus on chewing herbivores	28
Making small molecules in plants: A chassis for synthetic biology‐based production of plant natural products	28
The multi‐omics basis of potato heterosis	28
GmMs1 encodes a kinesin‐like protein essential for male fertility in soybean (Glycine max L.)	28
The transcription factors TLR1 and TLR2 negatively regulate trichome density and artemisinin levels in Artemisia annua	27
Mediator tail module subunits MED16 and MED25 differentially regulate abscisic acid signaling in Arabidopsis	27
Pathogen‐informed breeding for crop disease resistance	27
Dynamic regulation of DNA methylation and histone modifications in response to abiotic stresses in plants	27
Balancing selection and wild gene pool contribute to resistance in global rice germplasm against planthopper	27
Brassinosteroid signaling positively regulates abscisic acid biosynthesis in response to chilling stress in tomato	27
Understandings and future challenges in soybean functional genomics and molecular breeding	26
RING finger protein RGLG1 and RGLG2 negatively modulate MAPKKK18 mediated drought stress tolerance in Arabidopsis	26
Roles of DEMETER in regulating DNA methylation in vegetative tissues and pathogen resistance	26
Phase separation in plants: New insights into cellular compartmentalization	26
The Larix kaempferi genome reveals new insights into wood properties	26
An in planta haploid induction system in Brassica napus	26
Rice SPL10 positively regulates trichome development through expression of HL6 and auxin‐related genes	26
The E3 ligase XBAT35 mediates thermoresponsive hypocotyl growth by targeting ELF3 for degradation in Arabidopsis	26
The RNA N6‐methyladenosine demethylase ALKBH9B modulates ABA responses in Arabidopsis	25
OsEDM2L mediates m6A of EAT1 transcript for proper alternative splicing and polyadenylation regulating rice tapetal degradation	25
Phytochrome B interacts with SWC6 and ARP6 to regulate H2A.Z deposition and photomorphogensis in Arabidopsis	25
High non‐photochemical quenching of VPZ transgenic potato plants limits CO2 assimilation under high light conditions and reduces tuber yield under fluctuating light	25
A global alfalfa diversity panel reveals genomic selection signatures in Chinese varieties and genomic associations with root development	25
Tangeretin inhibits fungal ferroptosis to suppress rice blast	25
The alleviation of ammonium toxicity in plants	24
Comprehensive identification of lysine 2‐hydroxyisobutyrylated proteins in Ustilaginoidea virens reveals the involvement of lysine 2‐hydroxyisobutyrylation in fungal virulence	24
A G‐type lectin receptor‐like kinase regulates the perception of oomycete apoplastic expansin‐like proteins	24
HY5‐HDA9 orchestrates the transcription of HsfA2 to modulate salt stress response in Arabidopsis	24
A molecular framework for signaling crosstalk between jasmonate and ethylene in anthocyanin biosynthesis, trichome development, and defenses against insect herbivores in Arabidopsis	24
Biofortification of iron and zinc in rice and wheat	23
A bZIP transcription factor (CiFD) regulates drought‐ and low‐temperature‐induced flowering by alternative splicing in citrus	23
Interaction of brassinosteroid and cytokinin promotes ovule initiation and increases seed number per silique in Arabidopsis	23
Innovations and stepwise evolution of CBFs/DREB1s and their regulatory networks in angiosperms	22
Optimized prime editing efficiently generates heritable mutations in maize	22
HY5 regulates light‐responsive transcription of microRNA163 to promote primary root elongation in Arabidopsis seedlings	22
GhKNL1 controls fiber elongation and secondary cell wall synthesis by repressing its downstream genes in cotton (Gossypium hirsutum)	22
A histone H3K27me3 reader cooperates with a family of PHD finger‐containing proteins to regulate flowering time in Arabidopsis	22
Methionine oxidation and reduction of the ethylene signaling component MaEIL9 are involved in banana fruit ripening	22
Cultivated hawthorn (Crataegus pinnatifida var. major) genome sheds light on the evolution of Maleae (apple tribe)	22
Enhanced production of seed oil with improved fatty acid composition by overexpressing NAD+‐dependent glycerol‐3‐phosphate dehydrogenase in soybean	22
Ascorbate peroxidase 1 confers resistance to southern corn leaf blight in maize	21
Induction of jasmonic acid biosynthetic genes inhibits Arabidopsis growth in response to low boron	21
Primary root and root hair development regulation by OsAUX4 and its participation in the phosphate starvation response	21
Benefiting others and self: Production of vitamins in plants	21
Spatial phylogenetics of the Chinese angiosperm flora provides insights into endemism and conservation	21
Reciprocal regulation between the negative regulator PP2CG1 phosphatase and the positive regulator OST1 kinase confers cold response in Arabidopsis	21
The brassinosteroid signaling component SlBZR1 promotes tomato fruit ripening and carotenoid accumulation	21
Base editing‐mediated targeted evolution of ACCase for herbicide‐resistant rice mutants	21
BAK1 plays contrasting roles in regulating abscisic acid‐induced stomatal closure and abscisic acid‐inhibited primary root growth in Arabidopsis	21
Genome‐scale angiosperm phylogenies based on nuclear, plastome, and mitochondrial datasets	20
CmNF‐YB8 affects drought resistance in chrysanthemum by altering stomatal status and leaf cuticle thickness	20
Creation of fragrant sorghum by CRISPR/Cas9	20
Overlapping functions of YDA and MAPKKK3/MAPKKK5 upstream of MPK3/MPK6 in plant immunity and growth/development	20

A novel protein complex that regulates active DNA demethylation in Arabidopsis	20
Genome‐wide profiling of circular RNAs, alternative splicing, and R‐loops in stem‐differentiating xylem of Populus trichocarpa	20
Understanding the regulation of cereal grain filling: The way forward	19
GIGANTEA orthologs, E2 members, redundantly determine photoperiodic flowering and yield in soybean	19
Origin and evolution of green plants in the light of key evolutionary events	19
The ScCas9++ variant expands the CRISPR toolbox for genome editing in plants	19
Cowpea NAC1/NAC2 transcription factors improve growth and tolerance to drought and heat in transgenic cowpea through combined activation of photosynthetic and antioxidant mechanisms	19
WHIRLY1 recruits the histone deacetylase HDA15 repressing leaf senescence and flowering in Arabidopsis	19
Comparative transcriptomics reveals hidden issues in the plant response to arthropod herbivores	19
A domesticated Harbinger transposase forms a complex with HDA6 and promotes histone H3 deacetylation at genes but not TEs in Arabidopsis	19
Insights into salvianolic acid B biosynthesis from chromosome‐scale assembly of the Salvia bowleyana genome	19
The Phytophthora effector Avh241 interacts with host NDR1‐like proteins to manipulate plant immunity	19
Light participates in the auxin‐dependent regulation of plant growth	19
The Arabidopsis NuA4 histone acetyltransferase complex is required for chlorophyll biosynthesis and photosynthesis	19
Designing salt stress‐resilient crops: Current progress and future challenges	19
Zinc‐finger protein GmZF351 improves both salt and drought stress tolerance in soybean	19
PagGRF12a interacts with PagGIF1b to regulate secondary xylem development through modulating PagXND1a expression in Populus alba × P. glandulosa	18
MALE STERILITY 3 encodes a plant homeodomain‐finger protein for male fertility in soybean	18
New insights into cell–cell communications during seed development in flowering plants	18
ZmMYC2s play important roles in maize responses to simulated herbivory and jasmonate	18
ERF4 interacts with and antagonizes TCP15 in regulating endoreduplication and cell growth in Arabidopsis	18
Novel Wx alleles generated by base editing for improvement of rice grain quality	18
ANNEXIN 8 negatively regulates RPW8.1‐mediated cell death and disease resistance in Arabidopsis	18
Quantitative disease resistance: Multifaceted players in plant defense	18
Genome‐wide distribution and functions of the AAE complex in epigenetic regulation in Arabidopsis	18
The rice Raf‐like MAPKKK OsILA1 confers broad‐spectrum resistance to bacterial blight by suppressing the OsMAPKK4–OsMAPK6 cascade	18
Exploring the diversity of plant proteome	17
Regulation of root secondary metabolites by partial root‐associated microbiotas under the shaping of licorice ecotypic differentiation in northwest China	17
Agronomical selection on loss‐of‐function of GIGANTEA simultaneously facilitates soybean salt tolerance and early maturity	17
A cost‐effective tsCUT&Tag method for profiling transcription factor binding landscape	17
PROTEIN S‐ACYL TRANSFERASE 13/16 modulate disease resistance by S‐acylation of the nucleotide binding, leucine‐rich repeat protein R5L1 in Arabidopsis	17
Climate‐influenced boreotropical survival and rampant introgressions explain the thriving of New World grapes in the north temperate zone	17
OpNAC1 transcription factor regulates the biosynthesis of the anticancer drug camptothecin by targeting loganic acid O‐methyltransferase in Ophiorrhiza pumila	17
An efficient CRISPR/Cas9 platform for targeted genome editing in rose (Rosa hybrida)	17
The RECEPTOR‐LIKE PROTEIN53 immune complex associates with LLG1 to positively regulate plant immunity	16
Diversity, structure and function of the coiled‐coil domains of plant NLR immune receptors	16
An updated classification for the hyper‐diverse genusCorydalis(Papaveraceae: Fumarioideae) based on phylogenomic and morphological evidence	16
Regulation of cytokinin biosynthesis using PtRD26pro‐IPT module improves drought tolerance through PtARR10‐PtYUC4/5‐mediated reactive oxygen species removal inPopulus	16
Interplay between ethylene and nitrogen nutrition: How ethylene orchestrates nitrogen responses in plants	16
Ribonuclease H‐like gene SMALL GRAIN2 regulates grain size in rice through brassinosteroid signaling pathway	16
Assembly dynamics of East Asian subtropical evergreen broadleaved forests: New insights from the dominant Fagaceae trees	16
HBI1‐TCP20 interaction positively regulates the CEPs‐mediated systemic nitrate acquisition	16
Time series canopy phenotyping enables the identification of genetic variants controlling dynamic phenotypes in soybean	16
MYB106 is a negative regulator and a substrate for CRL3BPM E3 ligase in regulating flowering time in Arabidopsis thaliana	16
Plant transfer RNA‐derived fragments: Biogenesis and functions	16
Plant‐based, adjuvant‐free, potent multivalent vaccines for avian influenza virus via Lactococcus surface display	15
Natural variation in the NAC transcription factor NONRIPENING contributes to melon fruit ripening	15
Young Leaf White Stripe encodes a P‐type PPR protein required for chloroplast development	15
GmJAZ3 interacts with GmRR18a and GmMYC2a to regulate seed traits in soybean	15
High‐quality Fagopyrum esculentum genome provides insights into the flavonoid accumulation among different tissues and self‐incompatibility	15
Sdr4 dominates pre‐harvest sprouting and facilitates adaptation to local climatic condition in Asian cultivated rice	15
LjaFGD: Lonicera japonica functional genomics database	14
OsMAPK6 phosphorylates a zinc finger protein OsLIC to promote downstream OsWRKY30 for rice resistance to bacterial blight and leaf streak	14
Mediator complex subunit MED25 physically interacts with DST to regulate spikelet number in rice	14
Engineering of triterpene metabolism and overexpression of the lignin biosynthesis gene PAL promotes ginsenoside Rg3 accumulation in ginseng plant chassis	14
Growth asymmetry precedes differential auxin response during apical hook initiation in Arabidopsis	14
The Puccinia striiformis effector Hasp98 facilitates pathogenicity by blocking the kinase activity of wheat TaMAPK4	14
TRAF proteins as key regulators of plant development and stress responses	14
Genomic convergence underlying high‐altitude adaptation in alpine plants	14
ZmGRAS11, transactivated by Opaque2, positively regulates kernel size in maize	14
RING‐box proteins regulate leaf senescence and stomatal closure via repression of ABA transporter gene ABCG40	14
Three‐dimensional reconstruction and comparison of vacuolar membranes in response to viral infection	14
The NAC transcription factor ONAC083 negatively regulates rice immunity against Magnaporthe oryzae by directly activating transcription of the RING‐H2 gene OsRFPH2‐6	14
A critical review on natural compounds interacting with the plant plasma membrane H+‐ATPase and their potential as biologicals in agriculture	14
DNA methylation dynamics during germline development	14
The SlWRKY57‐SlVQ21/SlVQ16 module regulates salt stress in tomato	14
GmFtsH25 overexpression increases soybean seed yield by enhancing photosynthesis and photosynthates	14
Brassinosteroid signaling regulates phosphate starvation‐induced malate secretion in plants	14
The miR157‐SPL‐CNR module acts upstream of bHLH101 to negatively regulate iron deficiency responses in tomato	13
The Phytophthora effector Avh94 manipulates host jasmonic acid signaling to promote infection	13
Plant virology in the 21st century in China: Recent advances and future directions	13
Molecular basis of CONSTANS oligomerization in FLOWERING LOCUS T activation	13
Transcription factor GLK1 promotes anthocyanin biosynthesis via an MBW complex‐dependent pathway in Arabidopsis thaliana	13
The basic helix‐loop‐helix transcription factor gene, OsbHLH38, plays a key role in controlling rice salt tolerance	13
The safflower MBW complex regulates HYSA accumulation through degradation by the E3 ligase CtBB1	13
Establishment of an efficient seed fluorescence reporter‐assisted CRISPR/Cas9 gene editing in maize	13
DIW1 encoding a clade I PP2C phosphatase negatively regulates drought tolerance by de‐phosphorylating TaSnRK1.1 in wheat	13
Rapid improvement of rice eating and cooking quality through gene editing toward glutelin as target	13
NF‐YCs modulate histone variant H2A.Z deposition to regulate photomorphogenic growth in Arabidopsis	13
Bracelet salt glands of the recretohalophyte Limonium bicolor: Distribution, morphology, and induction	12
PagERF81 regulates lignin biosynthesis and xylem cell differentiation in poplar	12
PP2A interacts with KATANIN to promote microtubule organization and conical cell morphogenesis	12
Circadian rhythms driving a fast‐paced root clock implicate species‐specific regulation in Medicago truncatula	12
REVEILLE 7 inhibits the expression of the circadian clock gene EARLY FLOWERING 4 to fine‐tune hypocotyl growth in response to warm temperatures	12
Brassinosteroids inhibit miRNA‐mediated translational repression by decreasing AGO1 on the endoplasmic reticulum	12
ANAC087 transcription factor positively regulates age‐dependent leaf senescence through modulating the expression of multiple target genes in Arabidopsis	12
GhAP1‐D3 positively regulates flowering time and early maturity with no yield and fiber quality penalties in upland cotton	12
The pentatricopeptide repeat protein EMB1270 interacts with CFM2 to splice specific group II introns in Arabidopsis chloroplasts	11
Egg cell‐secreted aspartic proteases ECS1/2 promote gamete attachment to prioritize the fertilization of egg cells over central cells in Arabidopsis	11
Active DNA demethylation in plants: 20 years of discovery and beyond	11
Quantitative proteomics reveals redox‐based functional regulation of photosynthesis under fluctuating light in plants	11
The role of senescence‐associated gene101 (PagSAG101a) in the regulation of secondary xylem formation in poplar	11
Geminiviral C2 proteins inhibit active autophagy to facilitate virus infection by impairing the interaction of ATG7 and ATG8	11
The Xanthomonas type III effector XopAP prevents stomatal closure by interfering with vacuolar acidification	11
AtNSF regulates leaf serration by modulating intracellular trafficking of PIN1 in Arabidopsis thaliana	11